Pedipalps are composed of six segments or articles. From the proximal end (where they are attached to the spider) to the distal, they are: the coxa, the trochanter, the femur, the short patella, the tibia, and the tarsus. In spiders, the coxae frequently have extensions called maxillae or gnathobases, which function as mouth parts with or without some contribution from the coxae of the anterior legs. The limbs themselves may be simple tactile organs outwardly resembling the legs, as in spiders, or chelate weapons (pincers) of great size, as in scorpions. The pedipalps of Solifugae are covered in setae, but have not been studied in detail.[1]
Comparative studies of pedipalpal morphology may suggest that leg-like pedipalps are primitive in arachnids. At present, the only reasonable alternative to this view[according to whom?] is to assume that xiphosurans reflect the morphology of the primitive arachnid pedipalp and to conclude that this appendage is primitively chelate. Pedipalps are traditionally thought to be homologous with mandibles in crustaceans and insects, although more recent studies (e.g. using Hox genes) suggest they are probably homologous with the crustacean second antennae.
Pedipalps of spiders have the same segmentation as the legs, but the tarsus is undivided, and the pretarsus has no lateral claws. Pedipalps contain sensitive chemical detectors and function as taste and smell organs, supplementing those on the legs.[2] In sexually-mature male spiders, the final segment of the pedipalp, the tarsus, develops a complicated structure (sometimes called the palpal bulb or palpal organ) that is used to transfer sperm to the female seminal receptacles during mating. The details of this structure vary considerably between different groups of spiders and are useful for identifying species.[3][4] The pedipalps are also used by male spiders in courtship displays, contributing to vibratory patterns in web-shaking, acoustic signals, or visual displays.[5]
The cymbium is a spoon-shaped structure located at the end of the spider pedipalp that supports the palpal organ.[3] The cymbium may also be used as a stridulatory organ in spider courtship.[6]
The embolus is a narrow whip-like or leaf-like extension of the palpal bulb.
Bokma, John (photographer) (2005-09-07). Brachypelma vagans (Mexican red-rump) (photographs). Veracruz, MX. — Several close-up photos of a tarantula creating a sperm web