^ abDe Vos, L.; and L. Seegers (1998). Seven new Orthochromis species (Teleostei: Cichlidae) from the Malagarasi, Luiche and Rugufu basins (Lake Tanganyika drainage), with notes on their reproductive biology. Ichthyological Exploration of Freshwaters 9: 371—420.
^Schwarzer; Swartz; Vreven; Snoeks; Cotterill; Misof; and Schliewen (2012). Repeated trans-watershed hybridization among haplochromine cichlids (Cichlidae) was triggered by Neogene landscape evolution. Proc. R. Soc. B. 279: 4389—4398. doi:10.1098/rspb.2012.1667