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Luciobarbus
	Luciobarbus bocagei
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Cyprinidae
Subfamily:	Barbinae
Genus:	Luciobarbus; Heckel, 1843
Type species
	Luciobarbus esocinus;
Species
	see text
Synonyms
	Bertinius Fang, 1943; Messinobarbus Bianco, 1998

Luciobarbus is a genus of ray-finned fishes in the family Cyprinidae. Its members are found in fresh and brackish waters of southern Europe, northern Africa, the wider Near East, the Aral and Caspian Seas, and rivers associated with these. Several species in the genus are threatened. Most species are fairly small to medium-sized cyprinids, but the genus also includes several members that can surpass 1 m (3.3 ft) in length and the largest, the mangar (L. esocinus) can reach 2.3 m (7.5 ft).^[3]

Systematics

The type species is Luciobarbus esocinus, for which the genus was established by Heckel in 1843. The scientific name essentially means "pike-barbel" (after the northern pike, Esox lucius).^[4]

Like many other cyprinids, the present genus was long included in Barbus. It appears to be a very close relative of the typical barbels – which include that genus type species, Barbus barbus –, and may well warrant inclusion in Barbus. Many modern authors prefer to consider it a subgenus instead. It is, moreover, not entirely clear what species to place in Luciobarbus if it is deemed valid. The IUCN argues for a rather inclusive circumscription. Nonwithstanding the taxonomy and systematics of this ill-defined assemblage, their closest living relative is probably Aulopyge huegelii.^[5]

Species

The species in this genus:^[3]

Luciobarbus albanicus (Steindachner, 1870) (Albanian barbel)
Luciobarbus barbulus Heckel, 1847
Luciobarbus biscarensis Boulenger, 1911
Luciobarbus bocagei (Steindachner, 1864) (Iberian barbel)
Luciobarbus brachycephalus (Kessler, 1872) (Aral barbel)
Luciobarbus callensis Valenciennes, 1842 (Algerian barb)
Luciobarbus capito (Güldenstädt, 1773) (Bulatmai barbel)
Luciobarbus caspius (L. S. Berg, 1914) (Caspian barbel)
Luciobarbus chelifensis Brahimi, Freyhof, Henrard & Libois, 2017
Luciobarbus comizo (Steindachner, 1864) (Comizo barbel)
Luciobarbus escherichii (Steindachner, 1897) (Ankara barbel)
Luciobarbus esocinus Heckel, 1843 (Mangar)
Luciobarbus graecus Steindachner, 1896
Luciobarbus graellsii (Steindachner, 1866)
Luciobarbus guercifensis Doadrio, Perea & Yahyahoui, 2016 (Guercif barbel)
Luciobarbus guiraonis (Steindachner, 1866)
Luciobarbus kersin (Heckel, 1843) (Kersin barbel)
Luciobarbus kottelati Turan, Ekmekçi, İlhan & Engin, 2008 (Menderes barbel)
Luciobarbus lanigarensis Brahimi, Libois, Henrard & Freyhof, 2018
Luciobarbus leptopogon G. H. W. Schimper, 1834
Luciobarbus longiceps Valenciennes, 1842 (Jordan barbel)
Luciobarbus lydianus (Boulenger, 1896) (Lydian barbel)
Luciobarbus maghrebensis Doadrio, Perea & Yahyaoui, 2015 (Maghreb barbel) ^[6]
Luciobarbus magniatlantis (Pellegrin, 1919)
Luciobarbus microcephalus (Almaça (pt), 1967)
Luciobarbus mursa (Güldenstädt, 1773) (Mursa)
Luciobarbus mystaceus (Pallas, 1814) (Euphrates barbel)
†Luciobarbus nasus Günther, 1874)
Luciobarbus pallaryi Pellegrin, 1919
Luciobarbus pectoralis (Heckel, 1843) (Heckel’s Orontes barbel)
Luciobarbus rabatensis Doadrio, Perea & Yahyaoui, 2015 (Rabat barbel) ^[6]
Luciobarbus rifensis Doadrio, Casal-Lopez & Yahyaoui, 2015 (Rifian barbel) ^[6]
Luciobarbus sclateri (Günther, 1868) (Andalusian barbel)
Luciobarbus setivimensis Valenciennes, 1842
Luciobarbus steindachneri (Almaça (pt), 1967)
Luciobarbus subquincunciatus (Günther, 1868) (Mesopotamian barbel)
Luciobarbus xanthopterus Heckel, 1843 (Yellowfin barbel)
Luciobarbus yahyaouii Doadrio, Casal-Lopez & Perea, 2016 (Yahyaoui barbel)
Luciobarbus zayanensis Doadrio, Casal-Lopez & Perea, 2016 (Zayan barbel)

References

^ Bailly, Nicolas (2014). "Luciobarbus Heckel, 1843". WoRMS. World Register of Marine Species. Retrieved 6 February 2024.
^ BRUSLÉ Jacques; QUIGNARD Jean-Pierre (2013). "Chapitre 21 Les Barbeaux". Biologie des poissons d'eau douce européens (in French) (2e ed.). Lavoisier. p. 296. ISBN 9782743064969.
^ ^a ^b Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Luciobarbus". FishBase. August 2015 version.
^ "Family CYPRINIDAE: Subfamily BARBINAE Bleeker 1859 (Barbels)". The ETYFish Project. 7 June 2023. Retrieved 23 January 2024.
^ de Graaf et al. (2007), Almodóvar et al. (2008), IUCN (2009)
^ ^a ^b ^c Casal-Lopez, M., Perea, S., Yahyaoui, A. & Doadrio, I. (2015): Taxonomic review of the genus Luciobarbus Heckel, 1843 (Actinopterygii, Cyprinidae) from Northwestern Morocco with the description of three new species. Graellsia, 71 (2): e027.

Almodóvar, Ana; Nicola, Graciela G. & Elvira, Benigno (2008): Natural hybridization of Barbus bocagei x Barbus comizo (Cyprinidae) in Tagus River basin, central Spain [English with French abstract]. Cybium 32(2): 99-102. PDF fulltext
de Graaf, Martin; Megens, Hendrik-Jan; Samallo, Johannis & Sibbing, Ferdinand A. (2007): Evolutionary origin of Lake Tana's (Ethiopia) small Barbus species: indications of rapid ecological divergence and speciation. Anim. Biol. 57(1): 39-48. doi:10.1163/157075607780002069 (HTML abstract)