Sauropoda is a clade of dinosaurs that consists of roughly 300 species of large, long-necked herbivores and includes the largest terrestrial animals ever to exist. The first sauropod species were named in 1842 by Richard Owen, though at the time, he regarded them as unusual crocodilians. Sauropoda was named in 1878 by Othniel Marsh.[1]
Scope
For historical value, this list includes every sauropod species that has been formally named, regardless of whether the species is currently considered to be valid. Invalid species (i.e., species that are regarded as dubious or as a junior synonym of another species) are given a darker gray background.
Phylogenetic relationships of near-sauropod sauropodomorphs according to McPhee and Choiniere 2018,[2] with Sauropoda sensu lato highlighted in pink and Sauropoda sensu stricto highlighted in red to show the difference between the two definitions
The precise definition of Sauropoda is disputed. One proposed dividing line is to consider as sauropods all species more closely related to derived sauropods than Melanorosaurus, one of the most sauropod-like taxa historically classified as a "prosauropod". The other is to exclude from Sauropoda any species more distantly related to derived sauropods than Vulcanodon, one of the earliest and most basal taxa historically accepted as a sauropod. This leaves an interval of species more closely related to derived sauropods than Melanorosaurus but not so closely as Vulcanodon that are sauropods under one definition but not the other. Many characteristics of sauropods, such as opisthocoelous cervical vertebrae and columnar limbs, evolved within this transitional region, so that species within it may exhibit a combination of characteristics regarded as sauropod-like or non-sauropod-like.
For the purposes of this list, species whose phylogenetic position may fall within the interval between Melanorosaurus and Vulcanodon, or whose status as a sauropod is otherwise regarded as uncertain, are listed separately from unambiguous sauropods.
The third section of this list covers species that were once classified as sauropods, but have been reidentified as belonging to other groups.
The fourth section of this list covers informally named sauropod species.
Unambiguous sauropods
The following is a list of every species of sauropod that has ever been formally named, regardless of whether it is currently considered valid, dubious, or synonymous with another species. Invalid species are given a darker gray background. It does not contain species that have been recovered as basal to Vulcanodon by phylogenetic analysis, which are covered in a separate list below. The list contains the following information:
Scientific name: The binomial name of the species, accompanied by a citation to the work in which the species was formally named.
Status: The taxonomic status of the species, listing whether the species is currently regarded as valid, a nomen dubium, or as synonymous with another species.
Authors: The list of people credited with naming the species.
Date named: The date on which the species name is formally regarded as having first been published. According to the International Code of Zoological Nomenclature, only print publications and online works registered in ZooBank are considered to have been formally published,[3] so the date given is the date recorded by ZooBank or the date of print publication, if different from the date of online publication.
Parent clade: The most specific clade that the species is uncontroversially regarded as belonging to. When sorted, this column will place the clades in phylogenetic order, not alphabetical.
Age: The geological age of the species in millions of years ago. Ranges generally indicate uncertainty, not the known duration of the species, so the range of ages listed does not imply that the species existed for the entire duration of that range. When a numerical age range is not directly available, it is converted from the date range of the reported chronostratigraphic units.
Location: A list of countries where fossils of the species have been found. In the case of invalid species, only where the type specimen was found.
Notes: Miscellaneous notes, including what the species was originally called, if its genus assignment has been changed.
Skeletal diagram: A diagram showing a hypothetical silhouette of the animal, which bones are known of the species, and a size comparison if possible. For valid species, this may be a composite of multiple specimens, but for invalid species, this shows only the type specimen of the species.
Galveosaurus herreroi and Galvesaurus herreroi are based on the same skeleton but named by different authors, who have disputed which name is legitimate.
The generic name Jingia was originally proposed but it was already in use for a genus of moth.[92] The replacement name, Jingiella, was published in April later that year.[93]
This section of the list contains sauropodomorphs that are more closely related to eusauropods than Melanorosaurus but not as closely as Vulcanodon, thus lying between the two proposed boundaries of Sauropoda, as well as sauropodomorphs of unspecified phylogenetic position whose status as sauropods is disputed.
This section of the list covers taxa that were at one point referred to Sauropoda, its historical synonyms Cetiosauria or Opisthocoelia, or referred to a taxon that is now regarded as a sauropod, such as the genus Cetiosaurus. However, it excludes sauropodomorphs that were referred to Sauropoda as a result of debate over where the boundary between sauropods and non-sauropod sauropodomorphs lie.
Sauvage initially interpreted this species, based on a single vertebra, as belonging to Cetiosaurus. In 1895, he reinterpreted it as belonging to the marine reptile Pliosaurus.[104]
Listed as a member of Cetiosauria by Osborn in 1898.[105] Has been synonymized with Camptosaurus dispar, but may be a chimera that includes crocodylian material.[106]
Originally described as a sauropod under the name Neosaurus missouriensis,[107] but the name Neosaurus was preoccupied so it was renamed Parrosaurus missouriensis. Reclassified as a species of Hypsibema in 1979 by Baird and Horner, who regarded the genus as belonging to Sauropoda.[108]
Originally described as a species of the sauropod Astrodon. In 1981, Galton reidentified it as belonging to the stegosaurid Dacentrurus.[109] In 2008, Maidment et al. concluded it lacked evidence of stegosaur affinities and should be regarded as an indeterminate dinosaur.[110]
This section of the list contains sauropods that have never been formally scientifically named, but have been referred to with an informal binomial name.
Name
Author
Date
Notes
"Amphicoelias brontodiplodocus"
"Nurosaurus qaganensis"
"Mamenchisaurus guangyuanensis"
Zhang
1981
Described by Zhang Suoping in her 1981 master's thesis as "Omeisaurus guangyuanensis".[111] Referred to Mamenchisaurus by Li and Cai in 1997.[112] Ouyang and Ye tentatively agreed with Li and Cai's assessment in 2002, but were unsure whether it represented a distinct species.[111] Based on fossils from the Upper Shaximiao Formation, including several juvenile specimens.
"Dachongosaurus yunnanensis"
"Lancanjiangosaurus cachuensis"
"Megacervixosaurus tibetensis"
"Microdontosaurus dayensis"
"Oshanosaurus youngi"
"Rutellum implicatum"
"Yunxianosaurus hubeinensis"
Footnotes
^In 2004, Upchurch, Barrett, and Dodson listed Cetiosaurus brevis as a nomen dubium.[4] In 2011, Upchurch, Mannion, and Barrett regarded Cetiosaurus brevis and Pelorosaurus conybeari to be objective synonyms and, based on both the caudal vertebrae associated with the name C. brevis and humerus associated with P. conybeari, regarded the taxon, which they called Pelorosaurus conybeari, to be a valid species.[8]
^Some authors have treated Cetiosaurus conybeari Melville, 1849 and Pelorosaurus conybeari Mantell, 1850 as distinct taxa, with the former being based on a series of four caudal vertebrae and the latter on a humerus.[11][12][13] Others have regarded Pelorosaurus conybeari as simply Cetiosaurus conybeari assigned to a new genus.[14][10][8] Among those who treated P. conybeari as having been named by Mantell and based on a humerus, Blows regarded P. conybeari as a nomen vanum[15] and Upchurch et al. treated it as a valid taxon.[4]
^According to Naish and Martill, "sauropod experts seem in universal agreement that ʻC. gigasʼ is a camarasauromorph".[26] Naish and Martill themselves regarded Chondrosteosaurus to be a camarasaurid;[27] in contrast, Upchurch, Mannion, and Barrett regarded the specimen to be a titanosauriform.[8]
^Both Marsh and Lydekker named a sauropod genus Titanosaurus in 1877. Because Lydekker's Titanosaurus indicus was published first, Marsh renamed Titanosaurus montanus to Atlantosaurus montanus later that year.[30]
^In 2015, Tschopp, Mateus, and Benson argued that D. longus was a nomen dubium.[29] In 2017, Carpenter disputed their claims regarding the status of D. longus[34]
^Originally described as Ischyrosaurus, without a species name, by John W. Hulke in 1874.[24] Edward Drinker Cope had already named a genus Ischyrosaurus. In 1888, Lydekker assigned it to Ornithopsis, giving the species name as Ornithopsis manseli, which he credited to an unpublished manuscript by Hulke.[38]
^Hatcher incorrectly believed that Haplocanthus had to be changed because it was too similar to Haplacanthus Agassiz, 1845.[42] Nonetheless, the name Haplocanthosaurus became widely used, and in 1991, the ICZN ruled to conserve the name Haplocanthosaurus.[43]
^Gigantosaurus had to be renamed because Seeley had already used the name for a different sauropod
^In 1988, He et al. regarded it as uncertain whether O. changshouensis belonged to Omeisaurus or Mamenchisaurus.[51] In 1996, Zhang and Chen referred O. changshouensis to Mamenchisaurus in 1996.[52] In 2019, Tan et al. used the name Omeisaurus changshouensis but regarded it as more like Mamenchisaurus than Omeisaurus[53]
^Kim formally named a genus Ultrasaurus two years before Jensen did, so Ultrasaurus mcintoshi was renamed Ultrasauros.
^Issue 10 of volume 30 of the Canadian Journal of Earth Sciences is dated October 1993, but was not actually available until February or March 1994[56]
^Exact date disputed. Article dated July 1, 2005, but copies apparently did not become available until December 18, 2005.[67]
^The issue is dated 2009, but the paleontologist Darren Naish reported that it was actually published in 2008.[75]
^Originally described as a diplodocid, and recovered as such by Tschopp et al. in 2015[29] and Tschopp and Mateus in 2017.[77] In contrast, recovered as a dicraeosaurid by Whitlock and Wilson Mantilla in 2020.[39]
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Further reading
McIntosh, J. S. (1990). "Sauropoda". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.). The Dinosauria (1 ed.). Berkeley: University of California Press. pp. 345–401.
Olshevsky, George (1991-10-24). "A revision of the parainfraclass Archosauria Cope, 1869, excluding the advanced Crocodylia". Mesozoic Meanderings. 2: 1–196.
Upchurch, Paul; Barrett, Paul M.; Dodson, Peter (2004). "Sauropoda". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.). The Dinosauria (2 ed.). Berkeley: University of California Press. pp. 259–322. ISBN0-520-24209-2.