List of carnivorans by population
This is a list of estimated global populations of Carnivora species. This list is not comprehensive, as not all carnivorans have had their numbers quantified.
List
Common name
Binomial name
Population
Status
Trend
Notes
Image
Amur Leopard
Panthera pardus orientalis
20-30 [ 1]
CR [ 1]
[ 1]
Numbers have fluctuated in recent censuses.[ 1]
Iriomote cat
Prionailurus bengalensis iriomotensis
100–109 [ 2]
CR [ 3]
[ 3]
A subspecies of the leopard cat living exclusively on the Japanese island of Iriomote. Population size is declining, and consists of a single subpopulation.[ 2]
Malabar large-spotted civet
Viverra civettina
250 [ 4]
CR [ 4]
[ 4]
Maximum estimate. Previously considered possibly extinct.[ 4]
Darwin's fox
Pseudalopex fulvipes
250 [ 5]
EN [ 5]
[ 5]
Maximum estimate for mature individuals.[ 5]
Black-footed ferret
Mustela nigripes
500 [ 6]
EN [ 6]
[ 6]
Almost became extinct in the 1980s.[ 6]
Red wolf
Canis rufus
300 [ 7]
CR [ 7]
[ 7]
Maximum estimate. No more than 50 mature individuals. Previously extinct in the wild .[ 7]
Saimaa ringed seal
Pusa hispida saimensis
380
EN
The most endangered pinniped.
Cozumel raccoon
Procyon pygmaeus
323–955 [ 8]
CR [ 8]
[ 8]
Mature individuals number less than 250. Populations can be severely affected by hurricanes.[ 8]
Ethiopian wolf
Canis simensis
500 [ 9]
EN [ 9]
[ 9]
One of the world's rarest canids , and Africa's most endangered carnivore.
Siberian tiger
Panthera tigris tigris
560 [ 10]
EN [ 10]
[ 10]
Siberian tigers have made a comeback from the brink of extinction due to the strict laws of the Russian government. Russian Authorities believe that the number of living Siberian tigers will reach 700 after four years.[ 10]
Asiatic lion
Panthera leo leo
674 [ 11]
EN [ 11]
[ 11]
The population is rapidly growing from 284 in 1990 to 674 in 2020 thanks to the conservation efforts of the government, people and many communities. It is only found in the Gir forest and some other parts of the state of Gujrat, India.[ 11]
Mediterranean monk seal
Monachus monachus
700 [ 12]
VU [ 12]
[ 12]
The second most endangered pinniped.[ 12]
Marine otter
Lontra felina
1,000 [ 13]
EN [ 13]
[ 13]
Maximum estimate from 1991; numbers have probably fallen.[ 13]
Giant otter
Pteronura brasiliensis
1,000–5,000 [ 14]
EN [ 14]
[ 14]
Iberian lynx
Lynx pardinus
1,100 [ 15]
VU [ 16]
[ 16]
According to European Union LIFE Nature conservation projects estimates population size tripled from 52 mature individuals in 2002 to 156 in 2012.[ 16]
Hawaiian monk seal
Neomonachus schauinslandi
1,400 [ 17]
EN [ 17]
[ 17]
Numbered 1448 in 1983.[ 17]
Giant panda
Ailuropoda melanoleuca
1,800 [ 18]
VU [ 18]
[ 18]
Maximum estimate for mature individuals.[ 18]
Bay cat
Pardofelis badia
2,200 [ 19]
EN [ 19]
[ 19]
Maximum estimate.[ 19]
Dhole
Cuon alpinus
2,500 [ 20]
EN [ 20]
[ 20]
Maximum estimate for mature individuals.[ 20]
Fossa
Cryptoprocta ferox
2,500 [ 21]
VU [ 21]
[ 21]
Maximum estimate.[ 21]
Andean mountain cat
Leopardus jacobita
2,500 [ 22]
EN [ 22]
[ 22]
Maximum estimate.[ 22]
Grandidier's mongoose
Galidictis grandidieri
2,650–3,540 [ 23]
EN [ 23]
[ 23]
Snow leopard
Panthera uncia
<8,000 [ 24]
VU [ 24]
[ 24]
Bengal Tiger
Panthera tigris tigris
2,967 [ 25]
EN [ 25]
[ 25]
the population is rapidly growing and it has been doubled from 1,411 to 2,967 in 12 years. According to the census report, the population has been risen 30% from 2,226 in 2014 to 2,967 in 2020 thanks to the strict laws of Indian government.[ 25]
Tiger
Panthera tigris
3,900-5,600 [ 26]
EN [ 26]
[ 26]
75% of the tigers in the world belongs to India. 13 countries including India, Bangladesh, Myanmmar, Bhutan, China, Russia have taken a misson to double the worldwide tiger population by the year 2022.[ 26]
Island fox
Urocyon littoralis
4,001 [ 27]
NT
[ 27]
Sunda clouded leopard
Neofelis diardi
4,500 [ 28]
VU [ 28]
[ 28]
Maximum estimate.[ 6]
African wild dog
Lycaon pictus
6,600 [ 29]
EN [ 29]
[ 29]
Cheetah
Acinonyx jubatus
7,100 [ 30]
VU [ 30]
[ 30]
Estimate is for mature individuals only. Number may reach as high as 10,000.[ 30]
Brown hyena
Parahyaena brunnea
8,000 [ 31]
NT [ 31]
[ 31]
Eurasian lynx
Lynx lynx
8,000 (Europe only)[ 32]
LC [ 32]
[ 32]
Marbled cat
Pardofelis marmorata
10,000 [ 33]
NT [ 33]
[ 33]
Maximum estimate.[ 33]
Rusty-spotted cat
Prionailurus rubiginosus
10,000 [ 34]
NT [ 34]
[ 34]
Clouded leopard
Neofelis nebulosa
10,000 [ 35]
VU [ 35]
[ 35]
Maximum estimate for mature individuals.[ 35]
Red panda
Ailurus fulgens
10,000 [ 36]
EN [ 36]
[ 36]
African golden cat
Caracal aurata
10,000 [ 37]
VU
[ 37]
Minimum estimate for mature individuals.[ 37]
Chinese mountain cat
F. bieti
10,000 [ 38]
VU [ 38]
[ 38]
Maximum estimate for mature individuals.[ 38]
Galápagos fur seal
Arctophoca galapagoensis
10,000–15,000 [ 39]
EN [ 39]
[ 39]
Down from 30,000–40,000 in 1978.[ 39]
New Zealand sea lion
Phocarctos hookeri
11,855 [ 40]
EN [ 40]
[ 40]
Juan Fernández fur seal
Arctophoca philippii
12,000 [ 41]
LC [ 41]
[ 41]
Estimate is from the early 1990s; numbers appear to be increasing.[ 41]
Australian sea lion
Neophoca cinerea
13,790 [ 42]
EN [ 42]
[ 42]
Striped hyena
Hyaena hyaena
14,000 [ 43]
NT [ 43]
[ 43]
|
Sechuran fox
Lycalopex sechurae
15,000 [ 44]
NT [ 44]
Unknown[ 44]
Maximum estimate for mature individuals.[ 44]
Guadalupe fur seal
Arctophoca townsendi
15,000–17,000 [ 45]
LC [ 45]
[ 45]
Numbered 200 – 500 in the 1950s.[ 45]
Kodkod
Leopardus guigna
18,000 [ 46]
VU [ 46]
[ 46]
Maximum estimate.[ 46]
Leopard seal
Hydrurga leptonyx
18,000-35,000 [ 47]
LC [ 47]
Unknown[ 47]
Sloth bear
Melursus ursinus
20,000 [ 48]
VU [ 48]
[ 48]
Maximum estimate.[ 48]
Lion
Panthera leo
20,000 [ 49]
VU [ 49]
[ 49]
Comprises two subspecies in Africa and Asia . The Asiatic lion numbers an estimated 650.[ 50]
Galápagos sea lion
Zalophus wollebaeki
20,000–40,000 [ 51]
EN [ 51]
[ 51]
Polar bear
Ursus maritimus
22,000–31,000 [ 52]
VU [ 52]
[ 52]
Maned wolf
Chrysocyon brachyurus
23,600 [ 53]
NT [ 53]
Unknown[ 53]
Spotted hyena
Crocuta crocuta
27,000–47,000 [ 54]
LC [ 54]
[ 54]
Tentative estimate.[ 54]
Asian black bear
Ursus thibetanus
50,000 [ 55]
VU [ 56]
[ 56]
Maximum estimate.[ 56]
Cougar
Puma concolor
50,000 [ 57]
LC [ 57]
[ 57]
Jaguar
Panthera onca
64,000 [ 58]
NT [ 58]
[ 58]
Leopard
Panthera pardus
75,000 [ 59]
VU [ 59]
[ 59]
Baikal seal
Pusa sibirica
80,000–100,000 [ 60]
LC [ 60]
[ 60]
Sea otter
Enhydra lutris
106,822 [ 61]
EN [ 61]
[ 61]
Bush dog
Speothos venaticus
110,000 [ 62]
NT [ 62]
[ 62]
Considered an overestimate.[ 62]
Caspian seal
Pusa caspica
111,000 [ 63]
EN [ 63]
[ 63]
Once numbered over 1,000,000.[ 63]
Ross seal
Ommatophoca rossii
130,000 [ 64]
LC [ 64]
Unknown[ 64]
Steller sea lion
Eumetopias jubatus
143,000 [ 65]
NT [ 65]
[ 65]
Divided into two subspecies: Western Steller (78,000) and Loughlin's Steller (65,000).[ 65]
Northern elephant seal
Mirounga angustirostris
171,000 [ 66]
LC [ 66]
[ 66]
Numbered 113,000 in 1991.[ 66]
California sea lion
Zalophus californianus
180,000 [ 67]
LC [ 67]
[ 67]
Brown bear
Ursus arctos
200,000 [ 68]
LC [ 68]
[ 68]
Minimum estimate. Russia has the largest number of brown bears, believed to exceed 100,000, while estimates in the U.S. are around 33,000 (most in Alaska ), Canada 25,000, and Europe (excluding Russia) 14,000.[ 68]
New Zealand fur seal
Arctophoca forsteri
200,000 [ 69]
LC [ 69]
[ 69]
Subantarctic fur seal
Arctophoca tropicalis
200,000 [ 70]
LC [ 70]
[ 70]
[ 70]
South American sea lion
Otaria flavescens
250,000 [ 71]
LC [ 71]
[ 71]
Minimum estimate.[ 71]
South American fur seal
Arctophoca australis
250,000–300,000 [ 72]
LC [ 72]
[ 72]
Grey wolf
Canis lupus
300,000 [ 73]
LC
Unknown
Weddell seal
Leptonychotes weddellii
300,000 [ 74]
LC [ 74]
Unknown[ 74]
[ 74]
Harbor seal
Phoca vitulina
315,000 [ 75]
LC [ 75]
Unknown[ 75]
Founded in the coastal regions of North America, Europe and Asia.[ 75]
Grey seal
Halichoerus grypus
316,000 [ 76]
LC [ 76]
[ 76]
Southern elephant seal
Mirounga leonina
325,000 [ 77]
LC [ 77]
Unknown[ 77]
Latest global estimates are from the mid-1990s.[ 77]
Hooded seal
Cystophora cristata
340,000 [ 78]
VU [ 78]
[ 78]
One population in the Northwest Atlantic and another in the Northeast. The latter population has declined 85-90% in the past 60 years for unknown reasons.[ 78]
Northern fur seal
Callorhinus ursinus
650,000 [ 79]
VU [ 79]
[ 79]
American black bear
Ursus americanus
950,000 [ 80]
LC [ 80]
[ 80]
Numbers are believed to be twice that of all other bears combined.[ 80]
Brown fur seal
Arctocephalus pusillus
1,060,000 [ 81]
LC [ 81]
[ 81]
Divided into two subspecies: Cape and Australian.[ 81]
Crabeater seal
Lobodon carcinophaga
4,000,000 [ 82]
LC [ 82]
Unknown[ 82]
[ 82]
Harp seal
Pagophilus groenlandicus
4,500,000 [ 83]
LC [ 83]
[ 83]
[ 83]
Red fox
Vulpes vulpes
Unknown [ 84]
LC
Estimates are uncertain because a coordinated worldwide census has never been conducted.[ 84]
Cat (domestic)
Felis catus
600,000,000 [ 85]
Domesticated
[ 85]
2007 estimate
Dog (domestic)
Canis familiaris
900,000,000 [ 86]
Domesticated
[ 86]
See also
References
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^ a b 環境省. イリオモテヤマネコ生息状況等総合調査(第4次)の結果について(お知らせ) [(Fourth) Survey of the State of the Iriomote Cat's Habitat: Regarding the Results (notice)] (in Japanese). Retrieved 10 June 2012 .
^ a b Von Arx, M. & Breitenmoser-Wursten, C. (2008). "Prionailurus bengalensis iriomotensis" . IUCN Red List of Threatened Species . 2008 . Retrieved 2014-09-12 .
^ a b c d Mudappa, D.; Helgen, K.; Nandini, R. (2016). "Viverra civettina " . IUCN Red List of Threatened Species . 2016 : e.T23036A45202281. doi :10.2305/IUCN.UK.2016-1.RLTS.T23036A45202281.en .
^ a b c d Cuarón, A.D., de Grammont, P.C., Vázquez-Domínguez, E., Valenzuela-Galván, D., García-Vasco, D., Reid, F. & Helgen, K. (2008). "Pseudalopex fulvipes" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d e Belant, J.; Biggins, D.; Garelle, D.; Griebel, R.G.; Hughes, J.P. (2015). "Mustela nigripes " . IUCN Red List of Threatened Species . 2015 : e.T14020A45200314. doi :10.2305/IUCN.UK.2015-4.RLTS.T14020A45200314.en .
^ a b c d Kelly, B.T., Beyer, A. & Phillips, M.K. (2008). "Canis rufus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Khan, J.A. & Mukherjee, S. (2008). "Procyon pygmaeus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 .
^ a b c Marino, J. & Sillero-Zubiri, C. (2011). "Canis simensis" . IUCN Red List of Threatened Species . 2011 . Retrieved 2012-11-12 .
^ a b c d "Panthera tigris altaica" . IUCN Red List of Threatened Species . 2011 . 2011.
^ a b c d "Panthera leo persica" . IUCN Red List of Threatened Species . 2020 . 2020. Retrieved 2020-08-07 .
^ a b c d Aguilar, A. & Lowry, L. (IUCN SSC Pinniped Specialist Group) (2008). "Monachus monachus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Alvarez, R. & Medina-Vogel, G. (2008). "Lontra felina" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 .
^ a b c Duplaix, N., Waldemarin, H.F., Groenedijk, J., Evangelista, E., Munis, M., Valesco, M. & Botello, J.C. (2008). "Pteronura brasiliensis" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ Lusa (2021). "Pela primeira vez em 20 anos, há mais de 1000 linces-ibéricos" . Público .
^ a b c Rodríguez, A. (2024). "Lynx pardinus " . IUCN Red List of Threatened Species . 2024 : e.T12520A218695618. Retrieved 18 July 2024 .
^ a b c d Lowry, L. & Aguilar, A. (IUCN SSC Pinniped Specialist Group) (2008). "Monachus schauinslandi" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Lü, Z, Wang, D. & Garshelis, D.L. (IUCN SSC Bear Specialist Group) (2008). "Ailuropoda melanoleuca" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Hearn, A., Sanderson, J., Ross, J., Wilting, A. & Sunarto, S. (2008). "Pardofelis badia" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-16 . {{cite journal }}
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^ a b c d Durbin, L.S., Hedges, S., Duckworth, J.W., Tyson, M., Lyenga, A. & Venkataraman, A. (IUCN SSC Canid Specialist Group - Dhole Working Group) (2008). "Cuon alpinus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Hawkins, A.F.A. & Dollar, L. (2008). "Cryptoprocta ferox" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 .
^ a b c d Acosta, G., Cossios, D., Lucherini, M. & Villalba, L. (2008). "Leopardus jacobita" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Hawkins, A.F.A. (2008). "Galidictis grandidieri" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 .
^ a b c Jackson, R., Mallon, D., McCarthy, T., Chundaway, R.A. & Habib, B (2021). "Panthera uncia" . IUCN Red List of Threatened Species . 2021 . Retrieved 2021-02-06 . taking the high and low values of the population ranges provided (Table 1 in the Supporting Information) a range-wide population of 7,367-7,884 is indicated. If the results of the 2014-2016 Bhutan Snow Leopard census are added in (79-112: Thinley et al. 2016), the global population would then be estimated at 7,446 to 7,996. {{cite journal }}
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^ a b c d "Panthera tigris tigris" . IUCN Red List of Threatened Species . 2020 . 2020. Retrieved 2020-08-07 .
^ a b c d Chundawat, R.S., Habib, B., Karanth, U., Kawanishi, K., Ahmad Khan, J., Lynam, T., Miquelle, D., Nyhus, P., Sunarto, S., Tilson, R. & Sonam Wang (2011). "Panthera tigris" . IUCN Red List of Threatened Species . 2011 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b Coonan, T., Ralls, K., Hudgens, B., Cypher, B. & Boser, C (IUCN SSC Canid Specialist Group - Island Fox Working Group) (2013). "Urocyon littoralis" . IUCN Red List of Threatened Species . 2013 . Retrieved 2019-10-24 . {{cite journal }}
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^ a b c Hearn, A., Sanderson, J., Ross, J., Wilting, A. & Sunarto, S. (2008). "Neofelis diardi" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Woodroffe, R. & Sillero-Zubiri, C. (2012). "Lycaon pictus" . IUCN Red List of Threatened Species . 2012 . Retrieved 2012-11-13 .
^ a b c d Durant, S., Marker, L., Purchase, N., Belbachir, F., Hunter, L., Packer, C., Breitenmoser-Wursten, C., Sogbohossou, E. & Bauer, H. (2008). "Acinonyx jubatus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c Wiesel, I., Maude, G., Scott, D. & Mills, G. (2008). "Hyaena brunnea" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Breitenmoser, U., Mallon, D.P., von Arx, M. & Breitenmoser-Wursten, C. (2008). "Lynx lynx" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Grassman, L., Sanderson, J., Hearn, A., Ross, J., Wilting, A., Sunarto, S., Khan, J.A. & Mukherjee, S. (2008). "Pardofelis marmorata" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Khan, J.A. & Mukherjee, S. (2008). "Prionailurus rubiginosus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 .
^ a b c d Anderson, J., Khan, J.A., Grassman, L. & Mallon, D.P. (2008). "Neofelis nebulosa" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Wang, X., Choudhury, A., Yonzon, P., Wozencraft, C. & Than Zaw (2008). "Ailurus fulgens" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c Henschel, P., Breitenmoser-Wursten, C.& Sogbohossou, E. (2008). "Caracal aurata" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Sanderson, J., Mallon, D.P. & Driscoll, C. (2010). "F. silvestris bieti" . IUCN Red List of Threatened Species . 2010 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Aurioles, D. & Trillmich, F. (IUCN SSC Pinniped Specialist Group) (2008). "Arctocephalus galapagoensis" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c Gales, N. (IUCN SSC Pinniped Specialist Group) (2008). "Phocarctos hookeri" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 .
^ a b c d Aurioles, D. & Trillmich, F. (IUCN SSC Pinniped Specialist Group) (2008). "Arctocephalus philippii" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c Goldsworthy, S. & Gales, N. (IUCN SSC Pinniped Specialist Group) (2008). "Neophoca cinerea" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Arumugam, R., Wagner, A. & Mills, G. (2008). "Hyaena hyaena" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Asa, C.S., Cossíos, E.D. & Williams, R. (2008). "Pseudalopex sechurae" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c d Aurioles, D. & Trillmich, F. (IUCN SSC Pinniped Specialist Group) (2008). "Arctocephalus townsendi" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Acosta, G. & Lucherini, M. (2015). "Hydrurga leptonyx" . IUCN Red List of Threatened Species . 2015 . Retrieved 2020-06-13 .
^ a b c Hückstädt, L. (IUCN SSC Pinniped Specialist Group) (2015). "Hydrurga leptonyx" . IUCN Red List of Threatened Species . 2015 . Retrieved 2020-06-13 . APIS surveys resulted in an estimate of 35,500
^ a b c d Garshelis, D.L., Ratnayeke S. & Chauhan, N.P.S. (IUCN SSC Bear Specialist Group) (2008). "Melursus ursinus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Bauer, H.; Packer, C.; Funston, P.F.; Henschel, P.; Nowell, K. (2016). "Panthera leo " . IUCN Red List of Threatened Species . 2016 .
^ Kaushik, H. (August 2, 2017). "Lion population roars to 650 in Gujarat forests" . The Times of India . Retrieved 2017-11-24 .
^ a b c Aurioles, D. & Trillmich, F. (IUCN SSC Pinniped Specialist Group) (2008). "Zalophus wollebaeki" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Schliebe, S., Wiig, Ø., Derocher, A. & Lunn, N. (IUCN SSC Polar Bear Specialist Group) (2008). "Ursus maritimus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-13 . {{cite journal }}
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^ a b c Rodden, M., Rodrigues, F. & Bestelmeyer, S. (2008). "Chrysocyon brachyurus" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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^ a b c d Honer, O., Holekamp, K.E. & Mills, G. (2008). "Crocuta crocuta" . IUCN Red List of Threatened Species . 2008 . Retrieved 2012-11-12 . {{cite journal }}
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