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Granofilosea
	Clathrulina elegans
Scientific classification
Domain:	Eukaryota
Clade:	Diaphoretickes
Clade:	SAR
Phylum:	Cercozoa
Subphylum:	Reticulofilosa
Class:	Granofilosea; Cavalier-Smith & Bass 2009
Orders
	Desmothoracida; Cryptofilida; Limnofilida; Leucodictyida;

Granofilosea is a class of cercozoan protists in the subphylum Reticulofilosa.^[2] Out of the three groups that were traditionally considered heliozoans: the heliomonads, gymnosphaerids and desmothoracids, the latter were recently grouped into this new class.

Phylogeny and taxonomy

Phylogeny

The following cladogram shows the results of a 2011 phylogenetic analysis:^[1]

Monadofilosa

Granofilosea

novel clade Gran-5

	Desmothoracida

	novel clade Gran-4

novel clades Gran-2 & 3

Cryptofilida

	novel clade Gran-1

	Nanofilidae

novel clade Gran-6

Taxonomy

Class Granofilosea Cavalier-Smith & Bass 2009^[3]^[4]
- Family ?Microgromiidae De Saedeleer 1934
  - Genus Apogromia de Saedeleer 1934
  - Genus Belaria de Saedeleer 1934
  - Genus Heterogromia de Saedeleer 1934
  - Genus Microgromia Hertwig & Lesser 1874
  - Genus Paralieberkuehnia de Saedeleer 1934
- Order ?Axomonadida [Exonucleoaxoplastiales]
  - Family Tetradimorphidae Febvre-Chevalier & Febvre 1984
    - Genus Tetradimorpha Hsiung 1927
- Order Desmothoracida Honigberg et al. 1964 [Cryptaxohelida; Desmothoraca Hertwig & Lesser 1874; Clathrulinida Starobogatov 1980]
  - Family Clathrulinidae Claus 1874
    - Genus Actinosphaeridium Zacharias 1893
    - Genus Cienkowskya Schaudinn 1896 non Regel & Rach 1859 non Solms 1867
    - Genus Clathrulina Cienkowski 1867
    - Genus Elaeorhanis Greeff 1873
    - Genus Hedriocystis Hertwig & Lesser 1874
    - Genus Penardiophrys Mikrjukov 2000
- Order Cryptofilida Cavalier-Smith & Bass 2009
  - Family Mesofilidae Cavalier-Smith & Bass 2009
    - Genus Mesofila Cavalier-Smith & Bass 2009
  - Family Nanofilidae Cavalier-Smith & Bass 2009
    - Genus Nanofila Bass et al. 2009
- Order Limnofilida Cavalier-Smith & Bass 2009
  - Family Limnofilidae Cavalier-Smith & Bass 2009
    - Genus Limnofila Cavalier-Smith & Bass 2009
- Order Leucodictyida Cavalier-Smith 1993 emend. 2003
  - Family Massisteriidae Cavalier-Smith 1993
    - Genus Massisteria Larsen & Patterson 1990
    - Genus Minimassisteria Arndt & Cavalier-Smith 2011
  - Family Leucodictyidae Cavalier-Smith 1993 [Reticulamoebidae]
    - Genus Leucodictyon Grell 1991
    - Genus Reticulamoeba Grell 1994
    - Genus Thalassomyxa Grell 1985

References

^ ^a ^b Bass D, Chao EE, Nikolaev S, Yabuki A, Ishida K, Berney C, Pakzad U, Wylezich C, Cavalier-Smith T (2009). "Phylogeny of novel naked Filose and Reticulose Cercozoa: Granofilosea cl. n. and Proteomyxidea revised". Protist. 160 (1): 75–109. doi:10.1016/j.protis.2008.07.002. PMID 18952499.
^ Cavalier-Smith, Thomas (2017). "Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences". Protoplasma. 255 (1): 297–357. doi:10.1007/s00709-017-1147-3. PMC 5756292. PMID 28875267.
^ Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M. (2015-04-29). "A Higher Level Classification of All Living Organisms". PLOS ONE. 10 (4): e0119248. Bibcode:2015PLoSO..1019248R. doi:10.1371/journal.pone.0119248. ISSN 1932-6203. PMC 4418965. PMID 25923521.
^ Adl, S.M.; Simpson, A.G.B.; Lane, C.E.; Lukeš, J.; Bass, D.; Bowser, S.S.; Brown, M.W.; Burki, F.; Dunthorn, M.; Hampl, V.; Heiss, A.; Hoppenrath, M.; Lara, E.; le Gall, L.; Lynn, D.H.; McManus, H.; Mitchell, E.A.D.; Mozley-Stanridge, S.E.; Parfrey, L.W.; Pawlowski, J.; Rueckert, S.; Shadwick, L.; Schoch, C.L.; Smirnov, A.; Spiegel, F.W. (September 2012), "The Revised Classification of Eukaryotes", Journal of Eukaryotic Microbiology, 59 (5): 429–514, doi:10.1111/j.1550-7408.2012.00644.x, PMC 3483872, PMID 23020233