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Eocypraeidae
	Shell of Sphaerocypraea incomparabilis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Superfamily:	Cypraeoidea
Family:	Eocypraeidae; F. A. Schilder, 1924
Genera
	See text

Eocypraeidae is a small family of large sea snails, marine gastropod molluscs in the superfamily Cypraeoidea, the cowries and cowry allies.^[1]

The family Eocypraeidae consists mainly of fossil species, except for two species: Sphaerocypraea incomparabilis Briano, 1993 (regarded as valid) and an as yet undescribed species found in the southern Atlantic.^[2]

In 1963, a fresh shell of what was subsequently named Sphaerocypraea incomparabilis was dredged up by a Soviet trawler in the Gulf of Aden. There are currently (2009) only six known specimens of this shell in various collections.^[3]

Taxonomy

The taxonomy of Eocypraeidae is not fully understood and has undergone several modifications.

2005 taxonomy

According to the taxonomy of Bouchet & Rocroi (2005) this family belonged to the tribe Eocypraeini in the subfamily Ovulinae within the family Ovulidae.

2007 taxonomy

Fehse (2007)^[4] has raised the tribe Eocypraeini to the status of the family Eocypraeidae, based on morphological research of the radulae, shell and animal morphology and molecular phylogeny research of the 16S ribosomal RNA gene.^[4]

2016 taxonomy

Dolin thought that Eocypraeidae was part of the subfamily Prionovolvinae Fehse, 2007 in the family Ovulidae.^[5]

This line of thought was followed by Reijnen & van der Meij (2019)^[6] and even for a while by the database WoRMS and Zvonareva et al., 2020,^[7] each relying on a previous author.

2021 taxonomy

In 2021 Fehse accepted the systematic position of Eocypraeidae as valid and more consistent. Still he urges for a complete revision of the Eocypraeidae as well as the fossil Cypraeidae based exclusively on the type shells.^[8]

Genera

Genera within the family Eocypraeidae include:

Eocypraeinae F. A. Schilder, 1924

† Allmoniella Dolin & Dockery, 2018
† Apiocypraea F. A. Schilder, 1927
- † Apiocypraea davidi (Fontannes, 1880)
- † Apiocypraea parvoastensis (Sacco, 1894)
- † Apiocypraea spatiosa Fehse, 2003
† Cypropterina de Gregorio, 1880
† Eocypraea Cossmann, 1903 It is extinct since the end of the Eocene. This is the type genus of the family.
† Eschatocypraea F. A. Schilder, 1966
† Grovesia Dolin & Ledon, 2002
† Oxycypraea F. A. Schilder, 1927
† Taviania Dolin & Pacaud, 2009
† Borystheniella Pacaud, 2018 (uncertain > taxon inquirendum, controversed status)

Sulcocypraeinae Schilder, 1932

† Gaskoinia Roberts, 1870 (assignmrnt uncertain)
† Luponovula Sacco, 1894 †
Sphaerocypraea F. A. Schilder, 1927
† Sulcocypraea Conrad, 1865
† Willungia Powell, 1938

Synonyms

Chimaeria Briano, 1993: synonym of Sphaerocypraea F. A. Schilder, 1927
† Marginocypraea Ingram, 1947: synonym of Sphaerocypraea F. A. Schilder, 1927

References

^ ^a ^b MolluscaBase eds. (2022). MolluscaBase. Eocypraeidae F. A. Schilder, 1924. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=878564 on 2022-06-19
^ Fehse, D (2001). "Beiträge zur Kenntnis der Ovulidae (Mollusca: Cypraeoidea). VIII. Einleitung zur Familie sowie Katalog, Taxonomie und Bibliographie und Bemerkungen zu verwandten Gruppen". Acta Conchyliorum. 5.
^ Conniff, Richard (August 2009). "Mad About Seashells". Smithsonian magazine. Archived from the original on 2010-02-22. Retrieved 2010-01-02.
^ ^a ^b Fehse D. (2007). "Contributions to the knowledge of the Ovulidae. XVI. The higher systematics". Spixiana 30(1): 121-125. PDF Archived 2012-03-02 at the Wayback Machine.
^ Dolin, L. & Aguerre, O. (2016): Les Cypraeidae et les Ovulidae (Mollusca : Caenogastropoda) du Cuisien (Yprésien moyen) du bassin de Paris (France). Cossmanniana, 18: 18-37, pls. 1-6.
^ Reijnen, B.T. & van der Meij, S.E.T. (2019): Systematics of the subfamily Aclyvolvinae (Caenogastropoda: Ovulidae) based on molecular and morphometric analyses. Journal of Molluscan Studies, 85: 336-347, text figs. 1-6, tabs. 1-2.
^ Zvonareva, S.S., Mekhova, E.S., Hoang, D.T., Nguyen, T.H.T., VO, H.T. & Fedosov, A.E. (2020): Diversity and relationships of shallow-water Ovulidae (Mollusca: Gastropoda) of Vietnam. Archiv für Molluskenkunde, 149 (2): 113-146, text figs. 1-16, tabs. 1-4.
^ Fehse, D. (2021). The systematic position of the Eocypraeidae F.A. Schilder 1924 Part 2 (Mollusca: Gastropoda: Cypraeoidea). Privately published, Dirk Fehse, Berlin. 1-46