Alexander Kovalevsky

Alexander Kovalevsky
Alexander Kovalevsky
Born7 November 1840
Vārkava, Vitebsk Governorate, Russian Empire
Died1901
St. Petersburg, Russian Empire
NationalityRussian Empire
Alma materUniversity of Heidelberg
Known forGastrulation
Scientific career
FieldsEmbryology

Alexander Onufrievich Kovalevsky (Russian: Алекса́ндр Ону́фриевич Ковале́вский; 7 November 1840 – 1901) was a Russian embryologist, who studied medicine at the University of Heidelberg and became professor at the University of St Petersburg.[1][2][3][4] He was the brother of the paleontologist Vladimir Kovalevsky, and the brother-in-law of the mathematician Sofya Kovalevskaya.

Discoveries

A. Lancelet (a chordate), B. Larval tunicate, C. Adult tunicate. Kovalevsky saw that the notochord (1) and gill slit (5) are features shared by tunicates and vertebrates.

Kowalevsky's family belonged to Ukrainian nobility.

He showed that all animals go through a period of gastrulation.[1][2][3][4]

Kovalevsky discovered that tunicates are not molluscs, but that their larval stage has a notochord and pharyngeal slits, like vertebrates. Further, these structures develop from the same germ layers in the embryo as the equivalent structures in vertebrates, so he argued that the tunicates should be grouped with the vertebrates as chordates. 19th-century zoology thus converted embryology into an evolutionary science, connecting phylogeny with homologies between the germ layers of embryos, foreshadowing evolutionary developmental biology.[5]

Antoine Fortuné Marion and Alexander Kovalevsky, founder and collaborator of the Annals of Natural History Museum of Marseille

Honors

He was elected on the 1st of May 1884 a Foreign Member of the Linnean Society of London.[6] The St. Petersburg Society of Naturalists annually awards the A.O. Kovalevsky Medal.

Bibliography

  • Kowalevsky, A (1901). "Les Hedylidés, étude anatomique". Zapiski Imperatorskoi Akademii Nauk. 12: 1–32.

References

  1. ^ a b Ereskovsky, Alexander (2012). "Introduction to the Kowalevsky medal issue" (PDF). Evolution & Development. 14 (1): 1–2. doi:10.1111/j.1525-142X.2011.00516.x. PMID 23016968. S2CID 46420506.
  2. ^ a b Fokin, Sergei I. (2012). "Life of Alexander Onufrievich Kowalevsky (1840–1901)". Evolution & Development. 14 (1): 3–8. doi:10.1111/j.1525-142X.2011.00517.x. PMID 23016969. S2CID 205674995.
  3. ^ a b Haas, L F (May 1999). "Alexander Onufrievich Kovalevsky (1840-1901)". J. Neurol. Neurosurg. Psychiatry. 66 (5): 680. doi:10.1136/jnnp.66.5.680. PMC 1736354. PMID 10209188.
  4. ^ a b Kyle, R A; Shampo M A (Sep 1982). "Alexander O. Kovalevsky". JAMA. 248 (11): 1336. doi:10.1001/jama.248.11.1336. PMID 7050433.
  5. ^ Gilbert, Scott F. (2003). "The morphogenesis of evolutionary developmental biology" (PDF). International Journal of Developmental Biology. 47 (7–8): 467–477. PMID 14756322.
  6. ^ Proceedings of the Linnean Society of London (Session 1883–1884). 1880. p. 11.

Read other articles:

Lepas Masa Lalu

Lepas Masa LaluAlbum studio karya RadjaDirilis17 Juli 2001Direkam27–28 April 2001 di 144 Studio, BandungGenre Pop rock Alternative rock Rock Power ballad Durasi42:05LabelPT Ritme Nuansa Baru (RNB) (2005) ONE Music Universal Music Indonesia GP RecordsKronologi Radja Lepas Masa Lalu (2001) Manusia Biasa (2003)Manusia Biasa2003 Singel dalam album Lepas Masa Lalu Biar Aku MenjagamuDirilis: Juli 2001 Lepas Masa Lalu merupakan album musik pertama karya Radja yang dirilis pada bulan Juli 2001 ...

 

 

1815 en science

Chronologies Données clés 1812 1813 1814  1815  1816 1817 1818Décennies :1780 1790 1800  1810  1820 1830 1840Siècles :XVIIe XVIIIe  XIXe  XXe XXIeMillénaires :-Ier Ier  IIe  IIIe Chronologies géographiques Afrique Afrique du Sud, Algérie, Angola, Bénin, Botswana, Burkina Faso, Burundi, Cameroun, Cap-Vert, République centrafricaine, Comores, République du Congo, République démocratique du Congo, Côte d'Ivoire, Djibouti, Égyp...

 

 

جائزة بلجيكا الكبرى 1981

جائزة بلجيكا الكبرى 1981 السباق 5 من أصل 15 في بطولة العالم لسباقات الفورمولا واحد موسم 1981 السلسلة بطولة العالم لسباقات فورمولا 1 موسم 1981  البلد بلجيكا  التاريخ 17 مايو 1981 مكان التنظيم بلجيكا طول المسار 4.011 كيلومتر (2.492 ميل) المسافة 216.594 كيلومتر، (134.585 ميل) حالة الطقس جاف أ�...

الجيش الإمبراطوري الروسي

الجيش الإمبراطوري الروسيРусская императорская армия شارة الجيش الإمبراطوري الروسي الدولة  الإمبراطورية الروسية الإنشاء 1636 (إنشاء أول سفينة)1696 (الإنشاء بأمر من بيتر الأول). الانحلال 14 سبتمبر 1917  النوع قوات برية وحرس الدور الدفاع عن الإمبراطورية الروسية وحماية القيص�...

 

 

Spačva (river)

River in CroatiaSpačvaLocationCountryCroatiaPhysical characteristicsMouth  • locationBosut • coordinates45°03′52″N 19°04′35″E / 45.0645°N 19.0765°E / 45.0645; 19.0765Length40.2 km (25.0 mi)Basin featuresProgressionBosut→ Sava→ Danube→ Black Sea View of Spačva Spačva is a small river in eastern Croatia. It rises in the swampy Spačva region, takes in river Breznica, and flows into the Bosut be...

 

 

ماتيا كيجمان

ماتيا كيجمان (بالصربية اللاتينية: Mateja Kežman)‏  معلومات شخصية الميلاد 12 أبريل 1979 (العمر 44 سنة)بلغراد الطول 1.81 م (5 قدم 11 1⁄2 بوصة) مركز اللعب مهاجم الجنسية صربيا  مسيرة الشباب سنوات فريق 1986–1996 نادي زيمون المسيرة الاحترافية1 سنوات فريق م. (هـ.) 1996–1997 FK Radnički Pirot...

Ammatour

Municipality in Mount Lebanon, LebanonAmmatour عمّاطورMunicipalityAmmatourAmmatourLocation in Lebanon. The surrounding district can be seen on the map.Coordinates: 33°38′33″N 35°36′31″E / 33.64250°N 35.60861°E / 33.64250; 35.60861Country LebanonGovernorateMount LebanonDistrictChoufHighest elevation1,050 m (3,440 ft)Lowest elevation800 m (2,600 ft)Time zone+2 • Summer (DST)+3 Ammatour (Arabic: عمّاط�...

 

 

Grace Bedell

Correspondent with Abraham Lincoln The topic of this article may not meet Wikipedia's notability guideline for biographies. Please help to demonstrate the notability of the topic by citing reliable secondary sources that are independent of the topic and provide significant coverage of it beyond a mere trivial mention. If notability cannot be shown, the article is likely to be merged, redirected, or deleted.Find sources: Grace Bedell – news · newspapers · books...

 

 

Mireia Lalaguna

Nama ini menggunakan cara penamaan Spanyol: nama keluarga pertama atau paternalnya adalah Lalaguna dan nama keluarga kedua atau maternalnya adalah Royo. Mireia LalagunaMireia pada tahun 2016LahirMireia Lalaguna Royo28 November 1992 (umur 31)Barcelona, SpanyolPekerjaan Model Tinggi1,77 m (5 ft 9+1⁄2 in)Pemenang kontes kecantikanGelarMiss Atlántico Internacional 2014Miss World Spain 2015Warna rambutPirangWarna mataBiruKompetisiutamaMiss Atlántico Internacional 2...

Sweethearts (1938 film)

1938 film by Robert Zigler Leonard, W. S. Van Dyke SweetheartsTheatrical release posterDirected byW.S. Van DykeWritten bybook: Fred de GresacHarry B. SmithRobert B. SmithScreenplay byAlan CampbellDorothy ParkerLaura PerelmanS.J. PerelmanBased onSweethearts (1913)by Fred de GresacHarry B. SmithRobert Bache Smith[1][2][3]Produced byHunt StrombergStarringJeanette MacDonaldNelson EddyFrank MorganCinematographyOliver T. MarshAllen M. DaveyEdited byRobert KernMusic byVictor ...

 

 

Modena Football Club 1945-1946

Questa voce sull'argomento stagioni delle società calcistiche italiane è solo un abbozzo. Contribuisci a migliorarla secondo le convenzioni di Wikipedia. Segui i suggerimenti del progetto di riferimento. Voce principale: Modena Football Club. ModenaStagione 1945-1946Sport calcio Squadra Modena Allenatore Alfredo Mazzoni Presidente Luigi Pignatti Morano Divisione Nazionale6º (Alta Italia) Miglior marcatoreCampionato: Brighenti (10) 1943-44 1946-47 Si invita a seguire il modello d...

 

 

ロバート・デ・ニーロ

ロバート・デ・ニーロRobert De Niro 2011年のデ・ニーロ生年月日 (1943-08-17) 1943年8月17日（80歳）出生地 アメリカ合衆国・ニューヨーク州ニューヨーク市身長 177 cm職業 俳優、映画監督、映画プロデューサージャンル 映画、テレビドラマ活動期間 1963年 -配偶者 ダイアン・アボット（1976年 - 1988年）グレイス・ハイタワー（1997年 - ）主な作品 『ミーン・ストリート』（1973年）...

Timeline of Laredo, Texas

Timeline of the history of Laredo, Texas, United States The following is a timeline of the history of the city of Laredo, Texas, USA. This is a dynamic list and may never be able to satisfy particular standards for completeness. You can help by adding missing items with reliable sources. 18th-19th centuries Part of a series on the History of Texas Timeline Pre-Columbian Texas Early Spanish explorations 1519–1543 French Texas 1684–1689 Spanish Texas 1690–1821 Mexican Texas 18...

 

 

Irish Chess Championship

National event In various classes The Irish Chess Championship is the national Championship of Ireland, currently run by the Irish Chess Union (ICU), the FIDE-recognised governing body for the game.[1] Below is the list of champions. The first champion was J.A. Porterfield Rynd, who won the Dublin Chess Congress 1865 No. 3 Tournament, reserved for amateurs, bona fide resident in Ireland for the 12 months prior to 1st September 1865.[2] The Irish Chess Association was founded i...

 

 

Ing Yoe Tan

Artikel ini sebatang kara, artinya tidak ada artikel lain yang memiliki pranala balik ke halaman ini.Bantulah menambah pranala ke artikel ini dari artikel yang berhubungan atau coba peralatan pencari pranala.Tag ini diberikan pada November 2022. Dalam nama Tionghoa ini, nama keluarganya adalah Tan. Ing Yoe TanLahir(1948-04-18)18 April 1948Den Haag, BelandaMeninggal10 April 2020(2020-04-10) (umur 71)Amsterdam, BelandaPartai politikPartai Buruh (PvdA) Ing Yoe Tan Hanzi tradisional: 陳英�...

大字 (数字)

この項目には、一部のコンピュータや閲覧ソフトで表示できない文字が含まれています（詳細）。 数字の大字（だいじ）は、漢数字の一種。通常用いる単純な字形の漢数字（小字）の代わりに同じ音の別の漢字を用いるものである。 概要 壱万円日本銀行券（「壱」が大字） 弐千円日本銀行券（「弐」が大字） 漢数字には「一」「二」「三」と続く小字と、「壱」「�...

 

 

Station Agent

Station AgentI tre protagonisti in una scena del filmTitolo originaleThe Station Agent Paese di produzioneStati Uniti d'America Anno2003 Durata88 min Generecommedia, drammatico RegiaTom McCarthy SoggettoTom McCarthy SceneggiaturaTom McCarthy Casa di produzioneSenArt Films, Next Wednesday Productions Distribuzione in italianoBuena Vista International Italia FotografiaOliver Bokelberg MontaggioTom McArdle MusicheStephen Trask Interpreti e personaggi Peter Dinklage: Finbar McBride Paul Benja...

 

 

西遊記大結局之仙履奇緣

本條目存在以下問題，請協助改善本條目或在討論頁針對議題發表看法。 此條目需要編修，以確保文法、用詞、语气、格式、標點等使用恰当。 (2013年8月6日)請按照校對指引，幫助编辑這個條目。（幫助、討論） 此條目剧情、虛構用語或人物介紹过长过细，需清理无关故事主轴的细节、用語和角色介紹。 (2020年10月6日)劇情、用語和人物介紹都只是用於了解故事主軸，輔助�...

The Michael J. Fox Show

American television sitcom (2013–2014) The Michael J. Fox ShowGenreSitcomCreated by Will Gluck Sam Laybourne Starring Michael J. Fox Betsy Brandt Wendell Pierce Katie Finneran Juliette Goglia Conor Romero Jack Gore Ana Nogueira ComposerJeff CardoniCountry of originUnited StatesOriginal languageEnglishNo. of seasons1No. of episodes22 (7 unaired in the U.S.)ProductionExecutive producers Alex Reid Richard Schwartz Sam Laybourne Michael J. Fox Will Gluck Camera setupSingle-cameraRunning time20 ...

 

 

Camber (aerodynamics)

Asymmetry between the two acting surfaces of an airfoil In aeronautics and aeronautical engineering, camber is the asymmetry between the two acting surfaces of an airfoil, with the top surface of a wing (or correspondingly the front surface of a propeller blade) commonly being more convex (positive camber). An airfoil that is not cambered is called a symmetric airfoil. The benefits of cambering were discovered and first utilized by George Cayley in the early 19th century.[1] Overview ...