^Wedmann، Sonja؛ Wappler، Torsten؛ Engel، Michael S. (2009). "Direct and indirect fossil records of megachilid bees from the Paleogene of Central Europe (Hymenoptera: Megachilidae)". Naturwissenschaften. ج. 96 ع. 6: 703–712. Bibcode:2009NW.....96..703W. DOI:10.1007/s00114-009-0525-x. PMID:19296064.